The first two sequences demonstrate the usual interaction between territorial males and non-estrus females. In the subsequently shown copulation the male has already intromitted, holds the female down and is thrusting intravaginally. The female bites the male in the skin of his neck. In the next sequence an entire copulatory sequence is shown from initial repeated mounts, to intromission, slow intravaginal thrusting to the end of the copulation when the female struggles free.